Verbena stricta

Verbena stricta
Hoary vervain on a mixed-grass prairie of Lacreek NWR
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Lamiales
Family: Verbenaceae
Genus: Verbena
Species:
V. stricta
Binomial name
Verbena stricta
Vent.

Verbena stricta, also known as hoary verbena[1][2], hoary vervain[2][3], tall vervain[2], or woolly verbena[2], is a North American wildflower. It was first described by Étienne Pierre Ventenat in his work, Description des plantes nouvelles[4].

Distribution and Habitat

V. stricta is native to the contiguous United States, Quebec, and Ontario.[1] It is a drought-resistant species that prefers dry, well-drained soils and ample sunlight[2][5]. It is mostly found in low fields, meadows, prairies, and along roadsides and other disturbed habitats[2][3][5].

Description

In ideal growing conditions, V. stricta can grow up to 2–4 feet (0.6–1.2 m)[3][5]. It blooms for six weeks from June to September[2].

Flower

The inflorescence is made up of a densely clustered panicle of purple-blue or pink flowers along a hairy spike that may range from 1 to 8 inches long[2][3]. However, a mutation that causes lessened amounts of delphinidin and the complete loss of petunidin and malvidin produces white flowers[6]. Flowers are not all open at the same time[2]. Each flower is zygomorphic, consisting of a short corolla, four stamens, a small, toothed calyx, and five petals with a fused base, forming a slight tubular shape[2][3][5]. The petal lobes are unequal in size and length, with the two lateral lobes being slightly larger, and the bottom lobe bearing a small notch[3].

Leaves

The leaves are green in color and grow in an opposite arrangement along the side of the plant.[3] Their edges are heavily serrate, and the shape is oval to obovate with a pointed tip.[5] The stem and leaves of this plant are pubescent and may appear silvery-gray due to the density of the white hair that covers them, especially along the underside[3][5]. Leaves lack a petiole.[3][5]

Fruit

As the plant ages, the spike atop the plant will elongate. It will eventually seed and encapsulate four tiny nuts per flower. These fruit are usually 0.08–0.12 inches (2–3 mm) in length when ripe.[3][2]

Biology

Verbena stricta is facultatively xenogamous[7]: it can self-seed, but often cross-pollinates with V. hastata and V. urticifolia[8][7]. Hybrids are common among this species, giving rise to the Verbena stricta complex[8].

V. stricta is also of significance as a pollinator flower. In the southeast, is a host plant for the common buckeye butterfly[2] and is frequently visited by many long-tongued bees[5]. A specialist associated with this genus is the Nebraska vervain calliopsis bee[9]. Its foliage is commonly eaten by grasshoppers[5] and the seeds are eaten by songbirds[5][3]

Cultivation

V. stricta is prized as an ornamental plant due to its showy inflorescence and low maintenance, but it may also be considered a weed[5].

Some sources purport that V. stricta may have medical benefits, but these claims are not scientifically backed. It gained the folk moniker of "feverweed" for its use against fever in the 1800s.[10]

References

  1. ^ a b NRCS. "Verbena stricta". PLANTS Database. United States Department of Agriculture (USDA).
  2. ^ a b c d e f g h i j k l "Verbena stricta (Hoary Verbena, Hoary Vervain, Tall Vervain, Woolly Verbena) | North Carolina Extension Gardener Plant Toolbox". plants.ces.ncsu.edu. Retrieved 2025-12-09.
  3. ^ a b c d e f g h i j k Chayka, Katy; Dziuk, Peter (2016). "Verbena stricta (Hoary Vervain)". Minnesota Wildflowers. Retrieved 2017-09-17.
  4. ^ Ventenat, É. P.; Cels, Jacques-Martin; Redouté, Henri Joseph (1799). Description des plantes nouvelles et peu connues : cultivées dans le jardin de J.M. Cels : avec figures. A Paris: De l'imprimerie de Crapelet. doi:10.5962/bhl.title.66285.
  5. ^ a b c d e f g h i j k "Hoary Vervain (Verbena stricta)". www.illinoiswildflowers.info. Retrieved 2025-12-09.
  6. ^ Chai, Shengyue; Yang, Jiaming; Zhang, Xiaofei; Shang, Xuwen; Lang, Lixin (2024-11-21). "Unraveling the Anthocyanin Regulatory Mechanisms of White Mutation in Verbena stricta by Integrative Transcriptome and Metabolome Analysis". Genes. 15 (12): 1496. doi:10.3390/genes15121496. ISSN 2073-4425. PMC 11675223. PMID 39766764.
  7. ^ a b Cruden, Robert; Baker, Kristina; Cullinan, Thomas; Disbrow, Karen; Douglas, Kelly; Erb, John; Kirsten, Kenneth; Malik, Mary; Turner, Elizabeth; Weier, Jonathon; Wilmot, Sherry (1990-01-01). "The Mating Systems and Pollination Biology of Three Species of Verbena (Verbenaceae)". Journal of the Iowa Academy of Science. 97 (4): 178–183. ISSN 0896-8381.
  8. ^ a b Barber, Susan C. (October 1982). "Taxonomic Studies in the Verbena stricta Complex (Verbenaceae)". Systematic Botany. 7 (4): 433–456. Bibcode:1982SysBo...7..433B. doi:10.2307/2418676. JSTOR 2418676.
  9. ^ "Calliopsis nebraskensis (Nebraska Vervain Calliopsis Bee)". Minnesota Department of Natural Resources. Archived from the original on 2025-07-12. Retrieved 2025-12-09.
  10. ^ Coffey, Timothy (1993). The history and folklore of North American wildflowers. New York, NY: Facts on File. ISBN 978-0-8160-2624-1.