Portal:Devonian


The Devonian Portal

A map of Earth as it appeared 390 million years ago during the Middle Devonian Epoch

The Devonian (/dəˈvni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.62 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.86 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician Period.

Fishes, especially jawed fish, reached substantial diversity during this time, leading the Devonian to be called the Age of Fishes. The armored placoderms began dominating almost every known aquatic environment. In the oceans, cartilaginous fishes such as primitive sharks became more numerous than in the Silurian and Late Ordovician. Tetrapodomorphs, which include the ancestors of all four-limbed vertebrates (i.e. tetrapods), began diverging from freshwater lobe-finned fish as their more robust and muscled pectoral and pelvic fins gradually evolved into forelimbs and hindlimbs, though they were not fully established for life on land until the Late Carboniferous. (Full article...)

Selected Devonian Article

Partially reconstructed D. terrelli skull and trunk armor (specimen CMNH 5768), Cleveland Museum of Natural History

Dunkleosteus is an extinct genus of large arthrodire ("jointed-neck") fish that existed during the Late Devonian period, about 382–358 million years ago. It was a pelagic fish inhabiting open waters, and one of the first vertebrate apex predators of any ecosystem. Fossils of Dunkleosteus have been found in the United States, Canada, Poland, Belgium, and Morocco.

Dunkleosteus consists of ten species, some of which are among the largest placoderms ("plate-skinned") to have ever lived: D. terrelli, D. belgicus, D. denisoni, D. marsaisi, D. magnificus, D. missouriensis, D. newberryi, D. amblyodoratus, D. raveri, and D. tuderensis. However, the validity of several of these species is unclear. The largest and best known species is D. terrelli, the type species.

Since body shape is not known, various methods of estimation put the living total length of the largest known specimen of D. terrelli between 4.1 to 10 m (13 to 33 ft) long and its weight around 1–4 t (1.1–4.4 short tons). However, current evidence suggests length of less than 5 metres (16 ft) was more likely, with the most recent and extensive studies on the body shape and size of D. terrelli producing estimated lengths of approximately 3.4 metres (11 ft) for typical adults and 4.1 metres (13 ft) for exceptionally large individuals of this species. It has been suggested that Dunkleosteus could quickly open and close its jaw, possibly creating suction like modern-day suction feeders, and had an extremely high bite force, but both claims have recently been questioned. (Full article...)

Selected Devonian land plant article

Bark fragment from Sigillaria mamillaris sp. Estonian Museum of Natural History, Tallinn, Estonia.

Sigillaria is a genus of extinct, spore-bearing, arborescent lycophyte, known from the Carboniferous and Permian periods. It is related to the more famous Lepidodendron, and more distantly to modern quillworts. (Full article...)

Selected Devonian formation

Windom shale member of the Moscow formation, a trilobite pygidium (Eldredgeops rana) is exposed but not clearly visible

The Windom shale or Windom member is a member of the Moscow formation in New York. It preserves fossils going back to the Givetian stage of the middle Devonian. The Windom shale is widely known for its high quality trilobite fossils. (Full article...)

Selected Devonian fish article

Life restoration of two Cheirolepis

Cheirolepis (from Greek: χείρ kheír, 'hand' and Greek: λεπίς lepis 'scale') is an extinct genus of marine and freshwater ray-finned fish that lived in the Devonian period of Europe and North America. It is the only genus yet known within the family Cheirolepididae and the order Cheirolepidiformes. It was among the most basal of the Devonian actinopterygians and is considered the first to possess the "standard" dermal cranial bones seen in later actinopterygians.

Cheirolepis was a predatory freshwater and estuarine animal about 55 centimetres (22 in) long. It had a streamlined body with small, triangular ganoid scales similar to those of the Acanthodii. These scales had a basic structure typical of many early osteichthyans, with a superficial of ganoine overlying dentine, and a basal plate of bone. Cheirolepis had well-developed fins which gave it speed and stability, and was probably an active predator. Based on the size of its eyes, it hunted by sight. Cheirolepiss jaws, lined with sharp teeth, could be opened very wide, allowing it to swallow prey two thirds of its own size. ('Full article...)

Selected Devonian invertebrate

"Bryozoa", from Ernst Haeckel's Kunstformen der Natur, 1904

Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres (164 in) long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. 5,869 living species of bryozoa are known. Originally all of the crown group Bryozoa were colonial, but as an adaptation to a mesopsammal (interstitial spaces in marine sand) life or to deep-sea habitats, secondarily solitary forms have since evolved. Solitary species have been described in four genera: Aethozooides, Aethozoon, Franzenella, and Monobryozoon, the latter having a statocyst-like organ with a supposed excretory function.

The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups were noted to be very different internally. The new group was given the name "Entoprocta", while the original Bryozoa were called "Ectoprocta". Disagreements about terminology persisted well into the 20th century, but "Bryozoa" is now the generally accepted term.

Colonies take a variety of forms, including fans, bushes and sheets. Single animals, called zooids, live throughout the colony and are not fully independent. These individuals can have unique and diverse functions. All colonies have "autozooids", which are responsible for feeding, excretion, and supplying nutrients to the colony through diverse channels. Some classes have specialist zooids like hatcheries for fertilized eggs, colonial defence structures, and root-like attachment structures. Cheilostomata is the most diverse order of bryozoan, possibly because its members have the widest range of specialist zooids. They have mineralized exoskeletons and form single-layered sheets which encrust over surfaces, and some colonies can creep very slowly by using spiny defensive zooids as legs. (Full article...)

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