List of arvicolines
Arvicolinae is a subfamily of mammals in the rodent family Cricetidae, which in turn is part of the Myomorpha suborder in the order Rodentia. Members of this subfamily are called arvicolines and include voles, lemmings, and muskrats. They are found in North America, Europe, and Asia, primarily in forests, shrublands, grasslands, and wetlands, though some species can be found in deserts and rocky areas. They range in size from the woodland vole, at 6 cm (2 in) plus a 1 cm (0.4 in) tail, to the muskrat, at 30 cm (12 in) plus a 25 cm (10 in) tail. Arvicolines generally eat grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation, though some also eat fungus and invertebrates. Almost no arvicolines have population estimates, but three species—the Lake Baikal mountain vole, Tarabundí vole, and Zempoaltépec vole—are categorized as endangered.
The 158 extant species of Arvicolinae are divided into 30 genera, with over a third of species, 57, in the Microtus genus. Several extinct prehistoric arvicoline species have been discovered, though due to ongoing research and discoveries, the exact number and categorization are not fixed.[1]
Conventions
| Conservation status | |
|---|---|
| EX | Extinct (0 species) |
| EW | Extinct in the wild (0 species) |
| CR | Critically endangered (0 species) |
| EN | Endangered (3 species) |
| VU | Vulnerable (5 species) |
| NT | Near threatened (11 species) |
| LC | Least concern (126 species) |
| Other categories | |
| DD | Data deficient (13 species) |
| NE | Not evaluated (0 species) |
The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the arvicoline's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
Arvicolinae is a subfamily of the rodent family Cricetidae consisting of 158 extant species in 30 genera. These genera range in size from 1 to 57 species. This does not include hybrid species or extinct prehistoric species.
Subfamily Arvicolinae
- Genus Alexandromys (grass voles): eleven species
- Genus Alticola (high mountain voles): twelve species
- Genus Arborimus (tree voles): three species
- Genus Arvicola (water voles): three species
- Genus Caryomys (brownish voles): two species
- Genus Chionomys (snow voles): eleven species
- Genus Clethrionomys (bank voles): five species
- Genus Craseomys (grey-sided voles): six species
- Genus Dicrostonyx (collared lemmings): eight species
- Genus Dinaromys (Balkan snow vole): one species
- Genus Ellobius (mole voles): five species
- Genus Eolagurus (steppe lemmings): two species
- Genus Eothenomys (Père David's voles): eight species
- Genus Hyperacrius (Kashmir voles): two species
- Genus Lagurus (steppe lemming): one species
- Genus Lasiopodomys (narrow-headed steppe voles): two species
- Genus Lemmiscus (sagebrush vole): one species
- Genus Lemmus (lemmings): five species
- Genus Microtus (meadow voles): 57 species
- Genus Myopus (wood lemming): one species
- Genus Neodon (mountain voles): six species
- Genus Neofiber (round-tailed muskrat): one species
- Genus Ondatra (muskrat): one species
- Genus Phaiomys (Blyth's vole): one species
- Genus Phenacomys (heather voles): two species
- Genus Proedromys (Duke of Bedford's voles): two species
- Genus Prometheomys (long-clawed mole vole): one species
- Genus Stenocranius (narrow-headed voles): two species
- Genus Synaptomys (bog lemmings): two species
- Genus Volemys (Szechuan voles): two species
Arvicolines
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Evorsk vole
|
A. evoronensis (Kovalskaya & Sokolov, 1980) |
Southeastern Russia | Size: 12–16 cm (5–6 in) long, plus 4–7 cm (2–3 in) tail[4] Habitat: Grassland[5] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Japanese grass vole
|
A. montebelli (A. Milne-Edwards, 1872) |
Japan | Size: 9–14 cm (4–6 in) long, plus 2–5 cm (1–2 in) tail[4] Habitat: Forest and inland wetlands[7] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Lacustrine vole
|
A. limnophilus (Büchner, 1889) |
Mongolia and China | Size: 8–12 cm (3–5 in) long, plus 2–5 cm (1–2 in) tail[4] Habitat: Shrubland, grassland, rocky areas, and desert[8] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Maximowicz's vole
|
A. maximowiczii (Schrenck, 1859) |
Eastern Asia | Size: 11–16 cm (4–6 in) long, plus 2–6 cm (1–2 in) tail[4] Habitat: Forest[9] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Middendorff's vole
|
A. middendorffii (Poliakov, 1881) |
East-central Russia | Size: 9–15 cm (4–6 in) long, plus 2–5 cm (1–2 in) tail[4] Habitat: Forest, grassland, and inland wetlands[10] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Mongolian vole
|
A. mongolicus (Radde, 1861) |
Eastern Asia | Size: 8–13 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[4] Habitat: Forest and shrubland[11] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Muya Valley vole
|
A. mujanensis (Orlov & Kovalskaya, 1978) |
Southeastern Russia | Size: 11–16 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[4] Habitat: Forest[12] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Reed vole
|
A. fortis (Büchner, 1889) |
Eastern Asia | Size: 11–18 cm (4–7 in) long, plus 3–8 cm (1–3 in) tail[4] Habitat: Inland wetlands[13] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Sakhalin vole
|
A. sachalinensis (Vassin, 1955) |
Eastern Russia | Size: 10–18 cm (4–7 in) long, plus 3–8 cm (1–3 in) tail[4] Habitat: Grassland and inland wetlands[14] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Taiwan vole
|
A. kikuchii (Kuroda, 1920) |
Taiwan | Size: 10–14 cm (4–6 in) long, plus 8–10 cm (3–4 in) tail[4] Habitat: Forest and grassland[15] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Tundra vole
|
A. oeconomus (Pallas, 1776) |
Europe, northern Asia, and northwestern North America |
Size: 7–15 cm (3–6 in) long, plus 2–6 cm (1–2 in) tail[4] Habitat: Forest, shrubland, grassland, and inland wetlands[16] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Central Kashmir vole
|
A. montosa (F. W. True, 1894) |
Northern Pakistan and northern India | Size: 10–13 cm (4–5 in) long, plus 4–7 cm (2–3 in) tail[17] Habitat: Forest and rocky areas[18] Diet: Stems and leaves[19] |
VU
|
| Flat-headed vole
|
A. strelzowi (Kaschtschenko, 1899) |
Central Asia | Size: 10–14 cm (4–6 in) long, plus 3–5 cm (1–2 in) tail[17] Habitat: Grassland[20] Diet: Stems and leaves[19] |
LC
|
| Gobi Altai mountain vole
|
A. barakshin Bannikov, 1947 |
Central Asia | Size: 9–13 cm (4–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[17] Habitat: Grassland[21] Diet: Stems and leaves[19] |
LC
|
| Lake Baikal mountain vole
|
A. olchonensis Litvinov, 1960 |
Southern Russia | Size: 10–13 cm (4–5 in) long, plus 3–4 cm (1–2 in) tail[17] Habitat: Rocky areas[22] Diet: Stems and leaves[19] |
EN
|
| Large-eared vole
|
A. macrotis (Radde, 1862) |
Central Asia | Size: 9–12 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[17] Habitat: Rocky areas[23] Diet: Stems and leaves[19] |
LC
|
| Lemming vole
|
A. lemminus (G. S. Miller, 1899) |
Eastern Russia | Size: 8–12 cm (3–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[17] Habitat: Rocky areas[24] Diet: Stems and leaves[19] |
LC
|
| Mongolian silver vole
|
A. semicanus (Allen, 1928) |
Central Asia | Size: 9–14 cm (4–6 in) long, plus 2–4 cm (1–2 in) tail[17] Habitat: Grassland[25] Diet: Stems and leaves[19] |
LC
|
| Royle's mountain vole
|
A. roylei (J. E. Gray, 1842) |
Northern India | Size: 9–12 cm (4–5 in) long, plus 2–5 cm (1–2 in) tail[17] Habitat: Forest, shrubland, grassland, and rocky areas[26] Diet: Stems and leaves[19] |
NT
|
| Silver mountain vole
|
A. argentatus (Sévertsov, 1879) |
Central Asia | Size: 8–12 cm (3–5 in) long, plus 3–7 cm (1–3 in) tail[17] Habitat: Shrubland, grassland, and rocky areas[27] Diet: Stems and leaves[19] |
LC
|
| Stolička's mountain vole
|
A. stoliczkanus (Blanford, 1875) |
China and Nepal | Size: 9–13 cm (4–5 in) long, plus 1–3 cm (0–1 in) tail[17] Habitat: Forest, shrubland, grassland, and rocky areas[28] Diet: Stems and leaves[19] |
LC
|
| Tuva silver vole
|
A. tuvinicus Ognev, 1950 |
Central Asia | Size: 9–13 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[17] Habitat: Rocky areas[29] Diet: Stems and leaves[19] |
LC
|
| White-tailed mountain vole
|
A. albicaudus (True, 1894) |
Northern India | Size: 10–11 cm (4 in) long, plus 2–4 cm (1–2 in) tail[17] Habitat: Rocky areas and shrubland[30] Diet: Stems and leaves[19] |
DD
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Red tree vole
|
A. longicaudus (True, 1890) |
Western United States | Size: 9–11 cm (4 in) long, plus 6–9 cm (2–4 in) tail[31] Habitat: Forest[32] Diet: Conifer needles[33] |
NT
|
| Sonoma tree vole
|
A. pomo Johnson & George, 1991 |
Western United States |
Size: 9–11 cm (4 in) long, plus 6–8 cm (2–3 in) tail[31] Habitat: Forest[34] Diet: Conifer needles[33] |
NT
|
| White-footed vole
|
A. albipes (Merriam, 1901) |
Western United States | Size: 10–11 cm (4 in) long, plus 6–8 cm (2–3 in) tail[35] Habitat: Forest and inland wetlands[36] Diet: Conifer needles[33] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| European water vole
|
A. amphibius (Linnaeus, 1758) |
Europe and Asia |
Size: 12–23 cm (5–9 in) long, plus 5–14 cm (2–6 in) tail[37] Habitat: Forest, grassland, and inland wetlands[38] Diet: Aquatic plants, herbs, grass, twigs, buds, roots, bulbs, and fruit[39] |
LC
|
| Italian water vole
|
A. italicus Savi, 1838 |
South-central Europe | Size: 16–20 cm (6–8 in) long, plus 8–11 cm (3–4 in) tail[37] Habitat: Forest and inland wetlands[40] Diet: Aquatic plants, herbs, grass, twigs, buds, roots, bulbs, and fruit[39] |
NT
|
| Southwestern water vole
|
A. sapidus Miller, 1908 |
Western Europe |
Size: 16–22 cm (6–9 in) long, plus 10–14 cm (4–6 in) tail[37] Habitat: Inland wetlands[41] Diet: Aquatic plants, herbs, grass, twigs, buds, roots, bulbs, and fruit[39] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Ganzu vole
|
C. eva (Thomas, 1911) |
Central China | Size: 8–11 cm (3–4 in) long, plus 4–6 cm (2 in) tail[17] Habitat: Forest[42] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Kolan vole
|
C. inez (Thomas, 1908) |
Central China | Size: 8–11 cm (3–4 in) long, plus 2–5 cm (1–2 in) tail[17] Habitat: Forest[43] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Caucasian snow vole
|
C. gud Satunin, 1909 |
West-central Asia | Size: 11–13 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail[37] Habitat: Rocky areas[44] Diet: Grass, herbs, and seeds[45] |
LC
|
| European snow vole
|
C. nivalis (Martins, 1842) |
Europe and western Asia | Size: 10–15 cm (4–6 in) long, plus 4–8 cm (2–3 in) tail[37] Habitat: Rocky areas[46] Diet: Grass, herbs, and seeds[45] |
LC
|
| Robert's snow vole
|
C. roberti (Thomas, 1906) |
West-central Asia | Size: 12–16 cm (5–6 in) long, plus 8–11 cm (3–4 in) tail[37] Habitat: Forest[47] Diet: Grass, herbs, and seeds[45] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Bank vole
|
C. glareolus (Schreber, 1780) |
Europe and western Asia |
Size: 8–14 cm (3–6 in) long, plus 3–8 cm (1–3 in) tail[17] Habitat: Forest and shrubland[48] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Northern red-backed vole
|
C. rutilus (Pallas, 1779) |
Northeastern Europe, northern Asia, and northwestern North America |
Size: 8–13 cm (3–5 in) long, plus 2–5 cm (1–2 in) tail[31] Habitat: Forest and grassland[50] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Southern red-backed vole
|
C. gapperi (Vigors, 1830) |
Canada and United States |
Size: 10–12 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[31] Habitat: Forest and rocky areas[51] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Tien Shan red-backed vole
|
C. centralis (Miller, 1906) |
Central Asia | Size: 10–12 cm (4–5 in) long, plus 4–6 cm (2 in) tail[17] Habitat: Forest[52] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Western red-backed vole
|
C. californicus (Merriam, 1890) |
Western United States |
Size: 8–11 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail[31] Habitat: Forest[53] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Grey red-backed vole
|
C. rufocanus (Sundevall, 1846) |
Northern Europe and northern Asia |
Size: 10–15 cm (4–6 in) long, plus 2–7 cm (1–3 in) tail[31] Habitat: Forest, shrubland, grassland, inland wetlands, and rocky areas[54] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Hokkaido red-backed vole
|
C. rex (Imaizumi, 1971) |
Japan and eastern Russia | Size: 11–15 cm (4–6 in) long, plus 4–7 cm (2–3 in) tail[31] Habitat: Forest, shrubland, and grassland[55] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Japanese red-backed vole
|
C. andersoni (Thomas, 1905) |
Japan | Size: 7–14 cm (3–6 in) long, plus 4–8 cm (2–3 in) tail[31] Habitat: Inland wetlands and rocky areas[56] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Royal vole
|
C. regulus Thomas, 1907 |
South Korea and North Korea | Size: 10–12 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail[31] Habitat: Forest, shrubland, grassland, and rocky areas[57] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Shansei vole
|
C. shanseius (Thomas, 1908) |
Central and eastern China | Size: 9–11 cm (4 in) long, plus 2–4 cm (1–2 in) tail[31] Habitat: Forest, shrubland, grassland, inland wetlands, and rocky areas[58] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Smith's vole
|
C. smithii (Thomas, 1905) |
Japan | Size: 7–12 cm (3–5 in) long, plus 3–5 cm (1–2 in) tail[31] Habitat: Forest[59] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[49] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Arctic lemming
|
D. torquatus (Pallas, 1778) |
Northern Russia | Size: 8–14 cm (3–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Shrubland, grassland, and inland wetlands[60] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Nelson's collared lemming
|
D. nelsoni Merriam, 1900 |
Western Alaska |
Size: 12–14 cm (5–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[31] Habitat: Grassland[62] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Northern collared lemming
|
D. groenlandicus (Traill, 1823) |
Northern North America |
Size: 11–16 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Grassland[63] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Ogilvie Mountains collared lemming
|
D. nunatakensis Youngman, 1967 |
Northwestern Canada |
Size: 11–12 cm (4–5 in) long, plus 1–2 cm (0.4–0.8 in) tail[31] Habitat: Grassland[64] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Richardson's collared lemming
|
D. richardsoni Merriam, 1900 |
North-central Canada |
Size: 10–14 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[31] Habitat: Shrubland, grassland, and inland wetlands[65] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Unalaska collared lemming
|
D. unalascensis Merriam, 1900 |
Southwestern Alaska | Size: Unknown[31] Habitat: Grassland[66] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
DD
|
| Ungava collared lemming
|
D. hudsonius (Pallas, 1778) |
Northeastern Canada |
Size: 11–16 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Grassland and rocky areas[67] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
LC
|
| Wrangel lemming
|
D. vinogradovi (Ogniov, 1948) |
Eastern Russia | Size: 11–16 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Shrubland and grassland[68] Diet: Fruit, flowers, grass, sedges, buds, twigs, and bark[61] |
DD
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Balkan snow vole
|
D. bogdanovi (Martino & Martino, 1922) |
Southern Europe |
Size: 13–16 cm (5–6 in) long, plus 8–11 cm (3–4 in) tail[37] Habitat: Rocky areas[69] Diet: Grass[70] |
VU
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Alai mole vole
|
E. alaicus Vorontsov, Lyapunova, Zakaryan, & Ivanov, 1969 |
Southern Kyrgyzstan | Size: 11–14 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[37] Habitat: Unknown[71] Diet: Bulbs, tubers, and other underground plant parts[72] |
DD
|
| Northern mole vole
|
E. talpinus (Pallas, 1770) |
Eastern Europe and west-central Asia | Size: 8–12 cm (3–5 in) long, plus 0.5–3 cm (0.2–1.2 in) tail[37] Habitat: Grassland[73] Diet: Bulbs, tubers, and other underground plant parts[72] |
LC
|
| Southern mole vole
|
E. fuscocapillus (Blyth, 1843) |
West-central Asia | Size: 9–15 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[37] Habitat: Grassland[74] Diet: Bulbs, tubers, and other underground plant parts[72] |
LC
|
| Transcaucasian mole vole
|
E. lutescens (Thomas, 1897) |
Western Asia | Size: 10–14 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[37] Habitat: Grassland[75] Diet: Bulbs, tubers, and other underground plant parts[72] |
LC
|
| Zaisan mole vole
|
E. tancrei Blasius, 1884 |
Central Asia | Size: 9–13 cm (4–5 in) long, plus 1–2 cm (0.4–0.8 in) tail[37] Habitat: Shrubland, grassland, and desert[76] Diet: Bulbs, tubers, and other underground plant parts[72] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Przewalski's steppe lemming
|
E. przewalskii (Büchner, 1889) |
Mongolia and northern China | Size: 12–13 cm (5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[37] Habitat: Shrubland and grassland[77] Diet: Bulbs, seeds, and roots[78] |
LC
|
| Yellow steppe lemming
|
E. luteus (Eversmann, 1840) |
Central Asia | Size: 13–16 cm (5–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[37] Habitat: Shrubland[79] Diet: Bulbs, seeds, and roots[78] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Chaotung vole
|
E. olitor (Thomas, 1911) |
Southern China | Size: 8–10 cm (3–4 in) long, plus 2–4 cm (1–2 in) tail[17] Habitat: Forest[80] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Kachin red-backed vole
|
E. cachinus (Thomas, 1921) |
Southern China |
Size: 8–14 cm (3–6 in) long, plus 3–7 cm (1–3 in) tail[17] Habitat: Forest[82] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Pratt's vole
|
E. chinensis (Thomas, 1891) |
Central China | Size: 10–14 cm (4–6 in) long, plus 5–8 cm (2–3 in) tail[37] Habitat: Forest and grassland[83] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Père David's vole
|
E. melanogaster (H. Milne-Edwards, 1871) |
Eastern Asia |
Size: 7–12 cm (3–5 in) long, plus 2–6 cm (1–2 in) tail[17] Habitat: Forest, shrubland, and grassland[84] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Southwest China vole
|
E. custos (Thomas, 1912) |
Southern China | Size: 7–11 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail[37] Habitat: Forest, shrubland, grassland, and rocky areas[85] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Ward's red-backed vole
|
E. wardi (Thomas, 1912) |
Southern China | Size: 9–11 cm (4 in) long, plus 4–6 cm (2 in) tail[37] Habitat: Forest, grassland, and rocky areas[86] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
NT
|
| Yulungshan vole
|
E. proditor Hinton, 1923 |
Southern China | Size: 10–12 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[37] Habitat: Grassland and rocky areas[87] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
DD
|
| Yunnan red-backed vole
|
E. miletus (Thomas, 1914) |
South-central China | Size: 8–14 cm (3–6 in) long, plus 3–7 cm (1–3 in) tail[17] Habitat: Forest[88] Diet: Tender vegetation, nuts, seeds, bark, lichens, fungus, and insects[81] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Murree vole
|
H. wynnei (Blanford, 1881) |
Northern Pakistan and western China | Size: 10–14 cm (4–6 in) long, plus 2–5 cm (1–2 in) tail[37] Habitat: Forest and grassland[89] Diet: Grass, stems, and roots[70] |
LC
|
| True's vole
|
H. fertilis (True, 1894) |
Northern Pakistan and western China | Size: 8–12 cm (3–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[37] Habitat: Forest, shrubland, and grassland[90] Diet: Grass, stems, and roots[70] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Steppe lemming
|
L. lagurus (Pallas, 1773) |
Eastern Europe and western, central, and eastern Asia | Size: 7–13 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[37] Habitat: Shrubland and grassland[91] Diet: Green vegetation, tubers, and bulbs[78] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Brandt's vole
|
L. brandtii (Radde, 1861) |
East-central Asia | Size: 10–15 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[4] Habitat: Shrubland and grassland[92] Diet: Grass, vegetation, and roots[45] |
LC
|
| Mandarin vole
|
L. mandarinus (A. Milne-Edwards, 1871) |
Eastern Asia | Size: 9–12 cm (4–5 in) long, plus 2–3 cm (1–1 in) tail[4] Habitat: Grassland[93] Diet: Grass, vegetation, and roots[45] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Sagebrush vole
|
L. curtatus (Cope, 1868) |
Western United States and western Canada |
Size: 9–12 cm (4–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[37] Habitat: Shrubland and grassland[94] Diet: Green vegetation[95] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Amur lemming
|
L. amurensis Vinogradov, 1924 |
Eastern Russia and northeastern China |
Size: 8–12 cm (3–5 in) long, plus 1–2 cm (0.4–0.8 in) tail[31] Habitat: Shrubland, grassland, and inland wetlands[96] Diet: Sedges, grass, bark, leaves, berries, lichens, and roots[97] |
LC
|
| Canadian lemming
|
L. trimucronatus (Richardson, 1825) |
Eastern Russia and western North America |
Size: 11–14 cm (4–6 in) long, plus 1–2 cm (0.4–0.8 in) tail[31] Habitat: Grassland[98] Diet: Sedges, grass, bark, leaves, berries, lichens, and roots[97] |
LC
|
| Norway lemming
|
L. lemmus (Linnaeus, 1758) |
Northern Europe and northwestern Russia |
Size: 11–15 cm (4–6 in) long, plus 1–2 cm (0.4–0.8 in) tail[31] Habitat: Shrubland, grassland, and inland wetlands[99] Diet: Sedges, grass, bark, leaves, berries, lichens, and roots[97] |
LC
|
| West Siberian lemming
|
L. sibiricus (Kerr, 1792) |
Northern Russia |
Size: 9–16 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[31] Habitat: Forest, shrubland, grassland, and inland wetlands[100] Diet: Sedges, grass, bark, leaves, berries, lichens, and roots[97] |
LC
|
| Wrangel Island lemming
|
L. portenkoi Tchernyavsky, 1967 |
Wrangel Island in northeastern Russia | Size: 9–16 cm (4–6 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[31] Habitat: Grassland[101] Diet: Sedges, grass, bark, leaves, berries, lichens, and roots[97] |
DD
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Afghan vole
|
M. afghanus Thomas, 1912 |
Central Asia | Size: 6–13 cm (2–5 in) long, plus 1–4 cm (0.4–1.6 in) tail[102] Habitat: Shrubland, grassland, and desert[103] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Alpine pine vole
|
M. multiplex (Fatio, 1905) |
Southern Europe | Size: 9–11 cm (4 in) long, plus 4–5 cm (2 in) tail[104] Habitat: Forest and grassland[105] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Altai vole
|
M. obscurus Eversmann, 1841 |
Southeastern Europe and western and central Asia | Size: 10–14 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[104] Habitat: Grassland[106] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Anatolian vole
|
M. anatolicus Kryštufek & Kefelioğlu, 2002 |
Turkey | Size: 10–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Shrubland[107] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Beach vole
|
M. breweri Baird, 1858 |
Muskeget Island in northeastern United States | Size: 10–14 cm (4–6 in) long, plus 3–7 cm (1–3 in) tail[102] Habitat: Grassland and coastal marine[108] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
VU
|
| Bucharian vole
|
M. bucharensis Vinogradov, 1930 |
Central Asia | Size: About 13 cm (5 in) long, plus about 4 cm (2 in) tail[102] Habitat: Shrubland[109] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Cabrera's vole
|
M. cabrerae Thomas, 1906 |
Spain and Portugal |
Size: 10–14 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[102] Habitat: Forest, grassland, and inland wetlands[110] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
NT
|
| Calabria pine vole
|
M. brachycercus (Lehmann, 1961) |
Southern Italy | Size: 8–10 cm (3–4 in) long, plus 2–3 cm (1–1 in) tail[104] Habitat: Shrubland and grassland[111] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| California vole
|
M. californicus (Peale, 1848) |
Western United States and western Mexico |
Size: 11–14 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[102] Habitat: Grassland, desert, and intertidal marine[112] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Caspian gray vole
|
M. mystacinus (Filippi, 1865) |
Iran | Size: 10–13 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail[104] Habitat: Forest, shrubland, and grassland[113] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common vole
|
M. arvalis (Pallas, 1778) |
Europe and western Russia |
Size: 9–12 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Forest, shrubland, and grassland[114] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Creeping vole
|
M. oregoni (Bachman, 1839) |
Western United States and southwestern Canada |
Size: 9–12 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[104] Habitat: Forest, shrubland, and grassland[115] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Daghestan pine vole
|
M. daghestanicus (Shidlovsky, 1919) |
West-central Asia | Size: 9–11 cm (4 in) long, plus 3–5 cm (1–2 in) tail[104] Habitat: Inland wetlands[116] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Doğramaci's vole
|
M. dogramacii Kefelioğlu & Kryštufek, 1999 |
Turkey | Size: 9–13 cm (4–5 in) long, plus 1–4 cm (0.4–1.6 in) tail[104] Habitat: Shrubland[117] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| East European grey vole
|
M. rossiaemeridionalis (Ognev, 1924) |
Eastern Europe and western Asia | Size: 10–13 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail[104] Habitat: Grassland and inland wetlands[118] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Eastern meadow vole
|
M. pennsylvanicus (Ord, 1815) |
Canada and United States |
Size: 10–14 cm (4–6 in) long, plus 3–7 cm (1–3 in) tail[102] Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[119] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| European pine vole
|
M. subterraneus (Selys, 1836) |
Europe and western Asia |
Size: 7–12 cm (3–5 in) long, plus 2–5 cm (1–2 in) tail[104] Habitat: Forest, shrubland, and rocky areas[120] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Felten's vole
|
M. felteni Malec & Storch, 1963 |
Southeastern Europe | Size: 8–11 cm (3–4 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Forest[121] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Gerbe's vole
|
M. gerbei (Gerbe, 1879) |
France and Spain | Size: 9–11 cm (4 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Forest, grassland, and rocky areas[122] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Gray-tailed vole
|
M. canicaudus Miller, 1897 |
Northwestern United States |
Size: 10–13 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[102] Habitat: Grassland[123] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Guatemalan vole
|
M. guatemalensis Merriam, 1898 |
Guatemala and southern Mexico | Size: 11–13 cm (4–5 in) long, plus 3–4 cm (1–2 in) tail[102] Habitat: Forest[124] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
NT
|
| Günther's vole
|
M. guentheri (Danford & Alston, 1880) |
Western Asia | Size: 10–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Shrubland and grassland[125] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Insular vole
|
M. abbreviatus Miller, 1899 |
Alaska |
Size: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail[102] Habitat: Grassland[126] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Jalapan pine vole
|
M. quasiater (Coues, 1874) |
Eastern Mexico | Size: 9–11 cm (4 in) long, plus 1–3 cm (0.4–1.2 in) tail[102] Habitat: Forest, grassland, and rocky areas[127] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
NT
|
| Juniper vole
|
M. yuldaschi (Sévertsov, 1879) |
Central Asia | Size: 6–12 cm (2–5 in) long, plus 2–5 cm (1–2 in) tail[102] Habitat: Forest, shrubland, and grassland[128] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Kerman vole
|
M. kermanensis de Roguin, 1988 |
Iran | Size: 11–16 cm (4–6 in) long, plus 4–6 cm (2 in) tail[104] Habitat: Unknown[129] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Liechtenstein's pine vole
|
M. liechtensteini (Wettstein, 1927) |
Central and eastern Europe | Size: 8–12 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Forest and grassland[130] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Long-tailed vole
|
M. longicaudus (Merriam, 1888) |
Western United States and western Canada |
Size: 10–14 cm (4–6 in) long, plus 4–9 cm (2–4 in) tail[104] Habitat: Forest, shrubland, grassland, and inland wetlands[131] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Lusitanian pine vole
|
M. lusitanicus (Gerbe, 1879) |
Southwestern Europe |
Size: 8–10 cm (3–4 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Forest[132] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Major's pine vole
|
M. majori (Thomas, 1906) |
Western Asia | Size: 9–12 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[104] Habitat: Forest, shrubland, grassland, and inland wetlands[133] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Mediterranean field vole
|
M. lavernedii Crespon, 1844 |
Western and central Europe | Size: 9–14 cm (4–6 in) long, plus 2–6 cm (1–2 in) tail[102] Habitat: Shrubland, grassland, and inland wetlands[134] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Mediterranean pine vole
|
M. duodecimcostatus (Selys, 1839) |
Southwestern Europe |
Size: 8–12 cm (3–5 in) long, plus 1–4 cm (0.4–1.6 in) tail[104] Habitat: Shrubland[135] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Mexican vole
|
M. mexicanus (Saussure, 1861) |
Mexico and southwestern United States | Size: 10–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[102] Habitat: Forest[136] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Montane vole
|
M. montanus (Peale, 1848) |
Western United States |
Size: 11–16 cm (4–6 in) long, plus 2–7 cm (1–3 in) tail[102] Habitat: Shrubland, grassland, and inland wetlands[137] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| North American water vole
|
M. richardsoni (Kay, 1842) |
Western United States and western Canada |
Size: 12–18 cm (5–7 in) long, plus 6–10 cm (2–4 in) tail[102] Habitat: Grassland and inland wetlands[138] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Paradox vole
|
M. paradoxus (Ognev & Heptner, 1928) |
Turkmenistan and northeast Iran | Size: 8–13 cm (3–5 in) long, plus 2–3 cm (1–1 in) tail[104] Habitat: Shrubland and grassland[139] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Persian vole
|
M. irani Thomas, 1921 |
Iran | Size: 8–13 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Grassland[140] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
VU
|
| Portuguese field vole
|
M. rozianus Bocage, 1865 |
Northern Portugal and northwestern Spain |
Size: About 10 cm (4 in) long, plus about 4 cm (2 in) tail[102] Habitat: Grassland and inland wetlands[141] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Prairie vole
|
M. ochrogaster (Wagner, 1843) |
South-central Canada and central United States |
Size: 10–14 cm (4–6 in) long, plus 2–5 cm (1–2 in) tail[102] Habitat: Savanna, shrubland, grassland, and inland wetlands[142] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Qazvin vole
|
M. qazvinensis Golenishchev, Sablina, Borodin, & Gerasimov, 2003 |
Iran | Size: 8–13 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Shrubland and grassland[143] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Rock vole
|
M. chrotorrhinus (Miller, 1894) |
Eastern Canada and eastern United States |
Size: 9–13 cm (4–5 in) long, plus 4–7 cm (2–3 in) tail[102] Habitat: Forest and rocky areas[144] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Savi's pine vole
|
M. savii (de Sélys-Longchamps, 1838) |
Central Europe | Size: 8–10 cm (3–4 in) long, plus 2–3 cm (1–1 in) tail[104] Habitat: Forest and shrubland[145] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Schelkovnikov's pine vole
|
M. schelkovnikovi Satunin, 1907 |
Azerbaijan and Iran | Size: 8–12 cm (3–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[104] Habitat: Forest[146] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Schidlovsky's vole
|
M. schidlovskii Argiropulo, 1933 |
Eastern Europe and Turkey | Size: 8–13 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Grassland[147] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Short-tailed field vole
|
M. agrestis (Linnaeus, 1761) |
Europe and northern Asia |
Size: 9–14 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[102] Habitat: Forest, shrubland, grassland, and inland wetlands[148] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Sicilian pine vole
|
M. nebrodensis Palumbo, 1868 |
Island of Sicily in Italy | Size: 9–11 cm (4 in) long, plus 1–3 cm (0–1 in) tail[104] Habitat: Shrubland and grassland[149] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Singing vole
|
M. miurus Osgood, 1901 |
Alaska and northwestern Canada |
Size: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail[4] Habitat: Grassland and inland wetlands[150] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Social vole
|
M. socialis (Pallas, 1773) |
Central and western Asia | Size: 9–12 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[104] Habitat: Grassland[151] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Taiga vole
|
M. xanthognathus (Leach, 1815) |
Alaska and northern Canada |
Size: 14–18 cm (6–7 in) long, plus 4–6 cm (2 in) tail[102] Habitat: Forest and inland wetlands[152] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Tarabundí vole
|
M. oaxacensis Goodwin, 1966 |
Southern Mexico | Size: 11–13 cm (4–5 in) long, plus 3–4 cm (1–2 in) tail[102] Habitat: Forest[153] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
EN
|
| Tatra pine vole
|
M. tatricus (Kratochvíl, 1952) |
Eastern Europe | Size: 8–12 cm (3–5 in) long, plus 3–5 cm (1–2 in) tail[104] Habitat: Forest and grassland[154] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Thomas's pine vole
|
M. thomasi Barrett-Hamilton, 1903 |
Southeastern Europe | Size: 7–12 cm (3–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[104] Habitat: Grassland[155] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Tien Shan vole
|
M. ilaeus Thomas, 1912 |
Central Asia | Size: 10–15 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[156] Habitat: Forest, shrubland, and grassland[157] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Townsend's vole
|
M. townsendii (Bachman, 1839) |
Southwestern Canada and western United States |
Size: 12–16 cm (5–6 in) long, plus 4–8 cm (2–3 in) tail[102] Habitat: Grassland and inland wetlands[158] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Transcaspian vole
|
M. transcaspicus Satunin, 1905 |
Western Asia | Size: 9–14 cm (4–6 in) long, plus 3–6 cm (1–2 in) tail[156] Habitat: Grassland[159] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Woodland vole
|
M. pinetorum (Conte, 1830) |
Eastern Canada and eastern United States |
Size: 6–11 cm (2–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[102] Habitat: Forest and inland wetlands[160] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Zempoaltépec vole
|
M. umbrosus Merriam, 1898 |
Southern Mexico | Size: 11–14 cm (4–6 in) long, plus 4–7 cm (2–3 in) tail[102] Habitat: Forest[161] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Wood lemming
|
M. schisticolor (Lilljeborg, 1844) |
Northern Europe and northern Asia | Size: 9–12 cm (4–5 in) long, plus 1–2 cm (0.4–0.8 in) tail[31] Habitat: Forest and inland wetlands[162] Diet: Moss, stems, and bark[163] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Chinese scrub vole
|
N. irene (Thomas, 1911) |
Central and southern China | Size: 8–11 cm (3–4 in) long, plus 2–4 cm (1–2 in) tail[4] Habitat: Shrubland and grassland[164] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Clarke's vole
|
M. clarkei (Hinton, 1923) |
Southern China |
Size: 10–14 cm (4–6 in) long, plus 6–7 cm (2–3 in) tail[4] Habitat: Forest and grassland[165] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Forrest's mountain vole
|
N. forresti Hinton, 1923 |
Southern China and northern Myanmar |
Size: 10–14 cm (4–6 in) long, plus 3–5 cm (1–2 in) tail[4] Habitat: Grassland and rocky areas[166] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Linzhi mountain vole
|
N. linzhiensis Liu, Sun, Liu, Wang, Guo, & Murphy, 2012 |
Southern China and northern Myanmar | Size: 9–12 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[4] Habitat: Unknown[167] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Plateau vole
|
N. fuscus Büchner, 1889 |
Central China | Size: 9–15 cm (4–6 in) long, plus 2–5 cm (1–2 in) tail[4] Habitat: Grassland[168] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Sikkim mountain vole
|
N. sikimensis (Horsfield, 1841) |
Southern China and northern Bhutan | Size: 9–13 cm (4–5 in) long, plus 3–6 cm (1–2 in) tail[4] Habitat: Forest and grassland[169] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Round-tailed muskrat
|
N. alleni True, 1884 |
Southeastern United States |
Size: 18–21 cm (7–8 in) long, plus 9–17 cm (4–7 in) tail[35] Habitat: Grassland and inland wetlands[170] Diet: Maidencane, as well as rushes, sedges, sawgrass, and mangrove bark[171] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Muskrat
|
O. zibethicus (Linnaeus, 1766) |
North America, Europe, and Asia |
Size: 27–30 cm (11–12 in) long, plus 20–25 cm (8–10 in) tail[35] Habitat: Inland wetlands[172] Diet: Cattails, bulrushes, grass, and other vegetation, as well as crabs, crayfish, mussels, and small fish[173] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Blyth's vole
|
P. leucurus (Blyth, 1863) |
Western China | Size: 9–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail[4] Habitat: Forest, grassland, rocky areas, and shrubland[174] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Eastern heather vole
|
P. ungava (Merriam, 1889) |
Canada |
Size: 8–12 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[35] Habitat: Forest and shrubland[175] Diet: Bark, buds, heaths, forbs, berries, and seeds[176] |
LC
|
| Western heather vole
|
P. intermedius (Merriam, 1889) |
Western Canada and northwestern United States |
Size: 10–12 cm (4–5 in) long, plus 2–5 cm (1–2 in) tail[35] Habitat: Forest, shrubland, and grassland[177] Diet: Bark, buds, heaths, forbs, berries, and seeds[176] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Duke of Bedford's vole
|
P. bedfordi Thomas, 1911 |
Central China | Size: 9–13 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail[4] Habitat: Forest[178] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
VU
|
| Liangshan vole
|
P. liangshanensis (Liu, Sun, Zeng, & Zhao, 2007) |
Central China | Size: 10–14 cm (4–6 in) long, plus 6–9 cm (2–4 in) tail[4] Habitat: Forest and grassland[179] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Long-clawed mole vole
|
P. schaposchnikowi Satunin, 1901 |
Western Asia | Size: 12–17 cm (5–7 in) long, plus 3–6 cm (1–2 in) tail[35] Habitat: Grassland[180] Diet: Green vegetation and roots[72] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Narrow-headed vole
|
S. gregalis Pallas, 1789 |
Northern Asia | Size: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail[4] Habitat: Grassland[181] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Radde's vole
|
S. raddei (Poljakov, 1881) |
Northeastern Mongolia and southeastern Russia | Size: 8–15 cm (3–6 in) long, plus 1–5 cm (0.4–2.0 in) tail[4] Habitat: Grassland[182] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Northern bog lemming
|
S. borealis (Richardson, 1828) |
Canada and northern United States |
Size: 9–12 cm (4–5 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Forest, shrubland, grassland, and inland wetlands[183] Diet: Green vegetation as well as slugs, snails, and other invertebrates[184] |
LC
|
| Southern bog lemming
|
S. cooperi Baird, 1857 |
Eastern Canada and northeastern United States |
Size: 10–14 cm (4–6 in) long, plus 1–3 cm (0.4–1.2 in) tail[31] Habitat: Forest, shrubland, grassland, and inland wetlands[185] Diet: Green vegetation as well as slugs, snails, and other invertebrates[184] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Marie's vole
|
V. musseri (Lawrence, 1982) |
Central China | Size: 9–13 cm (4–5 in) long, plus 4–8 cm (2–3 in) tail[37] Habitat: Grassland and rocky areas[186] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
DD
|
| Szechuan vole
|
V. millicens (Thomas, 1911) |
Central China | Size: 8–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail[37] Habitat: Forest[187] Diet: Grass, leaves, twigs, bulbs, tubers, seeds, nuts, and other vegetation[6] |
NT
|
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- ^ a b Smith, A. T.; Johnston, C. H. (2016). "Myodes shanseius". IUCN Red List of Threatened Species. 2016 e.T7805A22373694. doi:10.2305/IUCN.UK.2016-2.RLTS.T7805A22373694.en.
- ^ a b Cassola, F. (2016). "Myodes smithii". IUCN Red List of Threatened Species. 2016 e.T16900A22372648. doi:10.2305/IUCN.UK.2016-2.RLTS.T16900A22372648.en.
- ^ a b Tsytsulina, K.; Formozov, N.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Dicrostonyx torquatus". IUCN Red List of Threatened Species. 2016 e.T6568A115082665. doi:10.2305/IUCN.UK.2016-3.RLTS.T6568A22332073.en.
- ^ a b c d e f g h Nowak, p. 1480
- ^ a b Cassola, F. (2016). "Dicrostonyx nelsoni". IUCN Red List of Threatened Species. 2016 e.T42620A22331765. doi:10.2305/IUCN.UK.2016-2.RLTS.T42620A22331765.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Dicrostonyx groenlandicus". IUCN Red List of Threatened Species. 2016 e.T42618A115195764. doi:10.2305/IUCN.UK.2016-3.RLTS.T42618A22331908.en.
- ^ a b Cassola, F. (2016). "Dicrostonyx nunatakensis". IUCN Red List of Threatened Species. 2016 e.T6567A22332006. doi:10.2305/IUCN.UK.2016-2.RLTS.T6567A22332006.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Dicrostonyx richardsoni". IUCN Red List of Threatened Species. 2016 e.T42622A115196066. doi:10.2305/IUCN.UK.2016-3.RLTS.T42622A22331579.en.
- ^ a b Garibaldi, A. (2019). "Dicrostonyx unalascensis". IUCN Red List of Threatened Species. 2019 e.T39974A22331447. doi:10.2305/IUCN.UK.2019-2.RLTS.T39974A22331447.en.
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- ^ a b Kryštufek, B.; Shenbrot, G. (2024). "Dinaromys bogdanovi". IUCN Red List of Threatened Species. 2024 e.T6607A221785055. doi:10.2305/IUCN.UK.2024-2.RLTS.T6607A221785055.en.
- ^ a b c Nowak, p. 1464
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- ^ a b c d e f g h Nowak, p. 1462
- ^ a b Smith, A. T. (2016). "Eothenomys cachinus". IUCN Red List of Threatened Species. 2016 e.T136507A22375635. doi:10.2305/IUCN.UK.2016-2.RLTS.T136507A22375635.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Eothenomys chinensis". IUCN Red List of Threatened Species. 2016 e.T7797A22375546. doi:10.2305/IUCN.UK.2016-1.RLTS.T7797A22375546.en.
- ^ a b Lunde, D.; Musser, G.; Molur, S. (2017) [errata version of 2016 assessment]. "Eothenomys melanogaster". IUCN Red List of Threatened Species. 2016 e.T7801A115086611. doi:10.2305/IUCN.UK.2016-3.RLTS.T7801A22375921.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Eothenomys wardi". IUCN Red List of Threatened Species. 2016 e.T136196A22375716. doi:10.2305/IUCN.UK.2016-2.RLTS.T136196A22375716.en.
- ^ a b Johnston, C.; Smith, A. T. (2019). "Eothenomys proditor". IUCN Red List of Threatened Species. 2019 e.T7803A22376123. doi:10.2305/IUCN.UK.2019-1.RLTS.T7803A22376123.en.
- ^ a b Smith, A. T.; Johnston, C. H. (2017) [errata version of 2016 assessment]. "Eothenomys miletus". IUCN Red List of Threatened Species. 2016 e.T136275A115205387. doi:10.2305/IUCN.UK.2016-3.RLTS.T136275A22376031.en.
- ^ a b Cassola, F. (2019). "Hyperacrius wynnei". IUCN Red List of Threatened Species. 2019 e.T10643A22353157. doi:10.2305/IUCN.UK.2019-1.RLTS.T10643A22353157.en.
- ^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Hyperacrius fertilis". IUCN Red List of Threatened Species. 2016 e.T10642A115098670. doi:10.2305/IUCN.UK.2016-3.RLTS.T10642A22353262.en.
- ^ a b Tsytsulina, K.; Formozov, N.; Zagorodnyuk, I.; Batsaikhan, N.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Lagurus lagurus". IUCN Red List of Threatened Species. 2016 e.T11179A115100995. doi:10.2305/IUCN.UK.2016-3.RLTS.T11179A22335535.en.
- ^ a b Avirmed, D.; Batsaikhan, N.; Tinnin, D. (2017) [errata version of 2016 assessment]. "Lasiopodomys brandtii". IUCN Red List of Threatened Species. 2016 e.T11340A115101423. doi:10.2305/IUCN.UK.2016-3.RLTS.T11340A22351917.en.
- ^ a b Batsaikhan, N.; Tsytsulina, K. (2017) [errata version of 2016 assessment]. "Lasiopodomys mandarinus". IUCN Red List of Threatened Species. 2016 e.T11342A115101561. doi:10.2305/IUCN.UK.2016-3.RLTS.T11342A22351819.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Lemmiscus curtatus". IUCN Red List of Threatened Species. 2016 e.T42624A115196202. doi:10.2305/IUCN.UK.2016-3.RLTS.T42624A22387210.en.
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- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Lemmus amurensis". IUCN Red List of Threatened Species. 2016 e.T11480A115102341. doi:10.2305/IUCN.UK.2016-3.RLTS.T11480A22335890.en.
- ^ a b c d e Nowak, p. 1481
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Lemmus trimucronatus". IUCN Red List of Threatened Species. 2016 e.T136712A115211700. doi:10.2305/IUCN.UK.2016-3.RLTS.T136712A22335983.en.
- ^ a b Amori, G. (2024). "Lemmus lemmus". IUCN Red List of Threatened Species. 2024 e.T11481A221786418. doi:10.2305/IUCN.UK.2024-2.RLTS.T11481A221786418.en.
- ^ a b Tsytsulina, K.; Formozov, N.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Lemmus sibiricus". IUCN Red List of Threatened Species. 2016 e.T11482A115102530. doi:10.2305/IUCN.UK.2016-3.RLTS.T11482A22336110.en.
- ^ a b Kennerley, R. (2019). "Lemmus portenkoi". IUCN Red List of Threatened Species. 2019 e.T136303A22336051. doi:10.2305/IUCN.UK.2019-1.RLTS.T136303A22336051.en.
- ^ a b c d e f g h i j k l m n o p q r s t u v w x Chernasky; Motis; Burgin, pp. 230–231
- ^ a b Molur, S. (2016). "Blanfordimys afghanus". IUCN Red List of Threatened Species. 2016 e.T2823A22383332. doi:10.2305/IUCN.UK.2016-2.RLTS.T2823A22383332.en.
- ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad Chernasky; Motis; Burgin, pp. 232–233
- ^ a b Amori, G. (2024). "Microtus multiplex". IUCN Red List of Threatened Species. 2024 e.T13490A221785322. doi:10.2305/IUCN.UK.2024-2.RLTS.T13490A221785322.en.
- ^ a b Kryštufek, B.; Amori, G. (2025). "Microtus obscurus". IUCN Red List of Threatened Species. 2025 e.T13450A22347022.
- ^ a b Dando, T.; Kennerley, R. (2021). "Microtus anatolicus". IUCN Red List of Threatened Species. 2021 e.T136237A137237409. doi:10.2305/IUCN.UK.2021-3.RLTS.T136237A137237409.en.
- ^ a b Roach, N. (2020). "Microtus breweri". IUCN Red List of Threatened Species. 2020 e.T13417A22349291. doi:10.2305/IUCN.UK.2020-2.RLTS.T13417A22349291.en.
- ^ a b Cassola, F. (2016). "Blanfordimys bucharensis". IUCN Red List of Threatened Species. 2016 e.T2824A22383414. doi:10.2305/IUCN.UK.2016-2.RLTS.T2824A22383414.en.
- ^ a b Amori, G. (2024). "Microtus cabrerae". IUCN Red List of Threatened Species. 2024 e.T13418A221784576. doi:10.2305/IUCN.UK.2024-2.RLTS.T13418A221784576.en.
- ^ a b Amori, G. (2024). "Microtus brachycercus". IUCN Red List of Threatened Species. 2024 e.T136536A221785859. doi:10.2305/IUCN.UK.2024-2.RLTS.T136536A221785859.en.
- ^ a b Álvarez-Castañeda, S. T.; Castro-Arellano, I.; Lacher, T.; Vázquez, E. (2016). "Microtus californicus". IUCN Red List of Threatened Species. 2016 e.T13427A22349460. doi:10.2305/IUCN.UK.2016-2.RLTS.T13427A22349460.en.
- ^ a b Mahmoudi, A.; Kryštufek, B.; Gazzard, A. (2025). "Microtus mystacinus". IUCN Red List of Threatened Species. 2025 e.T268982813A268982860. doi:10.2305/IUCN.UK.2025-1.RLTS.T268982813A268982860.en.
- ^ a b Amori, G. (2024). "Microtus arvalis". IUCN Red List of Threatened Species. 2024 e.T221687974A221337633. doi:10.2305/IUCN.UK.2024-2.RLTS.T221687974A221337633.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus oregoni". IUCN Red List of Threatened Species. 2016 e.T42632A115197199. doi:10.2305/IUCN.UK.2016-3.RLTS.T42632A22347079.en.
- ^ a b Kryštufek, B.; Bukhnikashvili, A.; Shenbrot, G. (2017) [errata version of 2016 assessment]. "Microtus daghestanicus". IUCN Red List of Threatened Species. 2016 e.T13428A115112364. doi:10.2305/IUCN.UK.2016-3.RLTS.T13428A22349378.en.
- ^ a b Kryštufek, B.; Yigit, N.; Kefelioğlu, H.; Sozen, M. (2017). "Microtus dogramacii". IUCN Red List of Threatened Species. 2017 e.T136535A22349286. doi:10.2305/IUCN.UK.2017-2.RLTS.T136535A22349286.en.
- ^ a b Kryštufek, B.; Mahmoundi, A. (2025). "Microtus rossiaemeridionalis". IUCN Red List of Threatened Species. 2025 e.T222269539A222269688. doi:10.2305/IUCN.UK.2025-1.RLTS.T222269539A222269688.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus pennsylvanicus". IUCN Red List of Threatened Species. 2016 e.T13452A115114123. doi:10.2305/IUCN.UK.2016-3.RLTS.T13452A22347596.en.
- ^ a b Hutterer, R.; Yigit, N.; Mitsainas, G.; Kryštufek, B.; Vohralík, V.; Zima, J.; Zagorodnyuk, I. (2021) [amended version of 2016 assessment]. "Microtus subterraneus". IUCN Red List of Threatened Species. 2021 e.T13489A197518317. doi:10.2305/IUCN.UK.2021-1.RLTS.T13489A197518317.en.
- ^ a b Amori, G. (2024). "Microtus felteni". IUCN Red List of Threatened Species. 2024 e.T13462A221785743. doi:10.2305/IUCN.UK.2024-2.RLTS.T13462A221785743.en.
- ^ a b Amori, G. (2024). "Microtus gerbii". IUCN Red List of Threatened Species. 2024 e.T39315A221785419. doi:10.2305/IUCN.UK.2024-2.RLTS.T39315A221785419.en.
- ^ a b Cassola, F. (2016). "Microtus canicaudus". IUCN Red List of Threatened Species. 2016 e.T42625A22348218. doi:10.2305/IUCN.UK.2016-2.RLTS.T42625A22348218.en.
- ^ a b Matson, J. (2020). "Microtus guatemalensis". IUCN Red List of Threatened Species. 2020 e.T13432A22350117. doi:10.2305/IUCN.UK.2020-2.RLTS.T13432A22350117.en.
- ^ a b Gazzard, A. (2023). "Microtus guentheri". IUCN Red List of Threatened Species. 2023 e.T221695743A221695823. doi:10.2305/IUCN.UK.2023-1.RLTS.T221695743A221695823.en.
- ^ a b Cassola, F. (2016). "Microtus abbreviatus". IUCN Red List of Threatened Species. 2016 e.T13425A22350031. doi:10.2305/IUCN.UK.2016-2.RLTS.T13425A22350031.en.
- ^ a b Álvarez-Castañeda, S. T.; Lacher, T.; Vázquez, E. (2019). "Microtus quasiater". IUCN Red List of Threatened Species. 2019 e.T13453A22348115. doi:10.2305/IUCN.UK.2019-3.RLTS.T13453A22348115.en.
- ^ a b Molur, S. (2023) [amended version of 2016 assessment]. "Microtus juldaschi". IUCN Red List of Threatened Species. 2023 e.T13436A239468807. doi:10.2305/IUCN.UK.2023-1.RLTS.T13436A239468807.en.
- ^ a b Kennerley, R.; Shenbrot, G. (2016). "Microtus kermanensis". IUCN Red List of Threatened Species. 2016 e.T13437A91356777. doi:10.2305/IUCN.UK.2016-3.RLTS.T13437A91356777.en.
- ^ a b Amori, G. (2024). "Microtus liechtensteini". IUCN Red List of Threatened Species. 2024 e.T221693652A221693787. doi:10.2305/IUCN.UK.2024-2.RLTS.T221693652A221693787.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus longicaudus". IUCN Red List of Threatened Species. 2016 e.T42627A115196586. doi:10.2305/IUCN.UK.2016-3.RLTS.T42627A22348744.en.
- ^ a b Amori, G. (2024). "Microtus lusitanicus". IUCN Red List of Threatened Species. 2024 e.T13494A221785647. doi:10.2305/IUCN.UK.2024-2.RLTS.T13494A221785647.en.
- ^ a b Kryštufek, B.; Shenbrot, G.; Sozen, M. (2016). "Microtus majori". IUCN Red List of Threatened Species. 2016 e.T136354A22346545. doi:10.2305/IUCN.UK.2016-2.RLTS.T136354A22346545.en.
- ^ a b Amori, G. (2024). "Microtus lavernedii". IUCN Red List of Threatened Species. 2024 e.T221686398A221686480. doi:10.2305/IUCN.UK.2024-2.RLTS.T221686398A221686480.en.
- ^ a b Amori, G. (2024). "Microtus duodecimcostatus". IUCN Red List of Threatened Species. 2024 e.T13493A221785542. doi:10.2305/IUCN.UK.2024-2.RLTS.T13493A221785542.en.
- ^ a b Álvarez-Castañeda, S. T.; Reid, F. (2017) [errata version of 2016 assessment]. "Microtus mexicanus". IUCN Red List of Threatened Species. 2016 e.T13443A115113184. doi:10.2305/IUCN.UK.2016-3.RLTS.T13443A22345679.en.
- ^ a b Cassola, F. (2016). "Microtus montanus". IUCN Red List of Threatened Species. 2016 e.T42630A22346732. doi:10.2305/IUCN.UK.2016-2.RLTS.T42630A22346732.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus richardsoni". IUCN Red List of Threatened Species. 2016 e.T42634A115197660. doi:10.2305/IUCN.UK.2016-3.RLTS.T42634A22345832.en.
- ^ a b Shenbrot, G.; Kryštufek, B. (2016). "Microtus paradoxus". IUCN Red List of Threatened Species. 2016 e.T136216A22351652. doi:10.2305/IUCN.UK.2016-2.RLTS.T136216A22351652.en.
- ^ a b Mahmoudi, A.; Kennerley, R. (2019). "Microtus irani". IUCN Red List of Threatened Species. 2019 e.T112465090A115589476. doi:10.2305/IUCN.UK.2019-1.RLTS.T112465090A115589476.en.
- ^ a b Amori, G. (2024). "Microtus rozianus". IUCN Red List of Threatened Species. 2024 e.T221686277A221686349. doi:10.2305/IUCN.UK.2024-2.RLTS.T221686277A221686349.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus ochrogaster". IUCN Red List of Threatened Species. 2016 e.T42631A115196932. doi:10.2305/IUCN.UK.2016-3.RLTS.T42631A22347375.en.
- ^ a b Kennerley, R. (2016). "Microtus qazvinensis". IUCN Red List of Threatened Species. 2016 e.T136565A22350870. doi:10.2305/IUCN.UK.2016-3.RLTS.T136565A22350870.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus chrotorrhinus". IUCN Red List of Threatened Species. 2016 e.T42626A115196387. doi:10.2305/IUCN.UK.2016-3.RLTS.T42626A22347958.en.
- ^ a b Amori, G. (2024). "Microtus savii". IUCN Red List of Threatened Species. 2024 e.T221713993A221715122. doi:10.2305/IUCN.UK.2024-2.RLTS.T221713993A221715122.en.
- ^ a b Kennerley, R. (2016). "Microtus schelkovnikovi". IUCN Red List of Threatened Species. 2016 e.T13456A22348673. doi:10.2305/IUCN.UK.2016-3.RLTS.T13456A22348673.en.
- ^ a b Shenbrot, G. (2016). "Microtus schidlovskii". IUCN Red List of Threatened Species. 2016 e.T136391A22349217. doi:10.2305/IUCN.UK.2016-2.RLTS.T136391A22349217.en.
- ^ a b Amori, G.; Gazzard, A. (2024). "Microtus agrestis". IUCN Red List of Threatened Species. 2024 e.T221684458A221685446. doi:10.2305/IUCN.UK.2024-2.RLTS.T221684458A221685446.en.
- ^ a b Amori, G. (2024). "Microtus nebrodensis". IUCN Red List of Threatened Species. 2024 e.T221713749A221713914. doi:10.2305/IUCN.UK.2024-2.RLTS.T221713749A221713914.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus miurus". IUCN Red List of Threatened Species. 2016 e.T42629A115196813. doi:10.2305/IUCN.UK.2016-3.RLTS.T42629A22346952.en.
- ^ a b Tsytsulina, K.; Kryštufek, B.; Yigit, N.; Bukhnikashvili, A.; Shenbrot, G. (2017) [errata version of 2016 assessment]. "Microtus socialis". IUCN Red List of Threatened Species. 2016 e.T13458A115114745. doi:10.2305/IUCN.UK.2016-3.RLTS.T13458A22348936.en.
- ^ a b Cassola, F. (2016). "Microtus xanthognathus". IUCN Red List of Threatened Species. 2016 e.T42628A22348489. doi:10.2305/IUCN.UK.2016-2.RLTS.T42628A22348489.en.
- ^ a b de Grammont, P. C.; Cuarón, A. D. (2018). "Microtus oaxacensis". IUCN Red List of Threatened Species. 2018 e.T13449A22346873. doi:10.2305/IUCN.UK.2018-1.RLTS.T13449A22346873.en.
- ^ a b Amori, G.; Shenbrot, G.; Kryštufek, B. (2024). "Microtus tatricus". IUCN Red List of Threatened Species. 2024 e.T13464A91985125. doi:10.2305/IUCN.UK.2024-2.RLTS.T13464A91985125.en.
- ^ a b Amori, G. (2024). "Microtus thomasi". IUCN Red List of Threatened Species. 2024 e.T13486A221784817. doi:10.2305/IUCN.UK.2024-2.RLTS.T13486A221784817.en.
- ^ a b Chernasky; Motis; Burgin, p. 234
- ^ a b Smith, A. T.; Johnston, C. H. (2016). "Microtus ilaeus". IUCN Red List of Threatened Species. 2016 e.T13438A22350742. doi:10.2305/IUCN.UK.2016-2.RLTS.T13438A22350742.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus townsendii". IUCN Red List of Threatened Species. 2016 e.T13487A115114983. doi:10.2305/IUCN.UK.2016-3.RLTS.T13487A22351048.en.
- ^ a b Shenbrot, G. (2016). "Microtus transcaspicus". IUCN Red List of Threatened Species. 2016 e.T13459A22349076. doi:10.2305/IUCN.UK.2016-2.RLTS.T13459A22349076.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Microtus pinetorum". IUCN Red List of Threatened Species. 2016 e.T42633A115197344. doi:10.2305/IUCN.UK.2016-3.RLTS.T42633A22346051.en.
- ^ a b de Grammont, P. C.; Cuarón, A. D. (2018). "Microtus umbrosus". IUCN Red List of Threatened Species. 2018 e.T42635A22346455. doi:10.2305/IUCN.UK.2018-1.RLTS.T42635A22346455.en.
- ^ a b Henttonen, H. (2017) [errata version of 2016 assessment]. "Myopus schisticolor". IUCN Red List of Threatened Species. 2016 e.T14104A115119850. doi:10.2305/IUCN.UK.2016-3.RLTS.T14104A22342119.en.
- ^ Nowak, p. 1482
- ^ a b Smith, A. T. (2016). "Neodon irene". IUCN Red List of Threatened Species. 2016 e.T13435A22330986. doi:10.2305/IUCN.UK.2016-2.RLTS.T13435A22330986.en.
- ^ a b Lunde, D.; Aplin, K. (2017) [errata version of 2016 assessment]. "Microtus clarkei". IUCN Red List of Threatened Species. 2016 e.T23041A115166908. doi:10.2305/IUCN.UK.2016-3.RLTS.T23041A22349944.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Neodon forresti". IUCN Red List of Threatened Species. 2016 e.T136554A22331098. doi:10.2305/IUCN.UK.2016-1.RLTS.T136554A22331098.en.
- ^ a b Gazzard, A. (2024). "Neodon linzhiensis". IUCN Red List of Threatened Species. 2024 e.T45959213A258447905. doi:10.2305/IUCN.UK.2024-2.RLTS.T45959213A258447905.en.
- ^ a b Johnston, C.; Smith, A. T. (2019). "Lasiopodomys fuscus". IUCN Red List of Threatened Species. 2019 e.T11341A22351731. doi:10.2305/IUCN.UK.2019-1.RLTS.T11341A22351731.en.
- ^ a b Molur, S. (2016). "Neodon sikimensis". IUCN Red List of Threatened Species. 2016 e.T13457A22330767. doi:10.2305/IUCN.UK.2016-2.RLTS.T13457A22330767.en.
- ^ a b Cassola, F. (2016). "Neofiber alleni". IUCN Red List of Threatened Species. 2016 e.T14520A22356567. doi:10.2305/IUCN.UK.2016-2.RLTS.T14520A22356567.en.
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- ^ a b Cassola, F. (2016). "Ondatra zibethicus". IUCN Red List of Threatened Species. 2016 e.T15324A22344525. doi:10.2305/IUCN.UK.2016-3.RLTS.T15324A22344525.en.
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- ^ a b Molur, S. (2016). "Phaiomys leucurus". IUCN Red List of Threatened Species. 2016 e.T13439A22333524. doi:10.2305/IUCN.UK.2016-2.RLTS.T13439A22333524.en.
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Phenacomys ungava". IUCN Red List of Threatened Species. 2016 e.T42637A115198018. doi:10.2305/IUCN.UK.2016-3.RLTS.T42637A22389703.en.
- ^ a b Nowak, p. 1466
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Phenacomys intermedius". IUCN Red List of Threatened Species. 2016 e.T42636A115197827. doi:10.2305/IUCN.UK.2016-3.RLTS.T42636A22389824.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Proedromys bedfordi". IUCN Red List of Threatened Species. 2016 e.T18305A22379426. doi:10.2305/IUCN.UK.2016-2.RLTS.T18305A22379426.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Proedromys liangshanensis". IUCN Red List of Threatened Species. 2016 e.T136459A22379536. doi:10.2305/IUCN.UK.2016-1.RLTS.T136459A22379536.en.
- ^ a b Kennerley, R. (2017). "Prometheomys schaposchnikowi". IUCN Red List of Threatened Species. 2017 e.T18339A22389618. doi:10.2305/IUCN.UK.2017-2.RLTS.T18339A22389618.en.
- ^ a b Amori, G.; Kennerley, R. (2024). "Stenocranius gregalis". IUCN Red List of Threatened Species. 2024 e.T221679815A221679871. doi:10.2305/IUCN.UK.2024-2.RLTS.T221679815A221679871.en.
- ^ a b Kennerley, R. (2024). "Stenocranius raddei". IUCN Red List of Threatened Species. 2024 e.T221679655A221679675. doi:10.2305/IUCN.UK.2024-2.RLTS.T221679655A221679675.en.
- ^ a b Cassola, F. (2017). "Synaptomys borealis". IUCN Red List of Threatened Species. 2017 e.T42638A22377185. doi:10.2305/IUCN.UK.2017-2.RLTS.T42638A22377185.en.
- ^ a b Nowak, p. 1483
- ^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Synaptomys cooperi". IUCN Red List of Threatened Species. 2016 e.T42639A115198182. doi:10.2305/IUCN.UK.2016-3.RLTS.T42639A22377358.en.
- ^ a b Johnston, C.; Smith, A. T. (2016). "Volemys musseri". IUCN Red List of Threatened Species. 2016 e.T23044A22387599. doi:10.2305/IUCN.UK.2016-1.RLTS.T23044A22387599.en.
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